Thought provoking article on pyramiding. Many know my reasons for wanting scientific explanations of husbandry protocols. The article does not promote anything...but gives an unbiased look at various research being done in the field. Simply food for thought.
Pyramiding - Big and Small Tortoise Rescue and Sanctuary
...the proliferation of the epidermis in scutes of the carapace and plastron in the tortoise (Testudo hermanni) corresponded with seasonal changes.
bigandsmalltortoise.org