I came across another great article. This one is titled "Influence of the calcium content of the diet offered to leopard tortoises (Geochelone pardalis)".
Just reading the abstract you can get the idea of how critical calcium supplementation is in captive tortoises.
I spoke with Danny quite awhile ago about supplementing and he suggested that tortoises might require A LOT more supplementing than is often recommended, and this study seems to confirm what he has said.
It is written specifically about leopard tortoises, but it can be applicable to other tortoise species.
http://africantortoise.com/leopard.pdf
Just reading the abstract you can get the idea of how critical calcium supplementation is in captive tortoises.
I spoke with Danny quite awhile ago about supplementing and he suggested that tortoises might require A LOT more supplementing than is often recommended, and this study seems to confirm what he has said.
It is written specifically about leopard tortoises, but it can be applicable to other tortoise species.
http://africantortoise.com/leopard.pdf