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I can't find the source where I read it (looking around for it) but I do remember a paper that said pyramiding was due to low humidity levels.

I've been lurking for a while so I might as well register so I can comment and post too. I've been keeping turtles for about 30 years (still have one of my first turtles). I also have 2 sulcatas, 4 redfoots and a large collection of box turtles. Look forward to participating and maybe...